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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
17.2
nTPM
HPA Brain:
48.5
nTPM
GTEx:
17.2
nTPM
FANTOM5:
52.9
Scaled Tags Per Million
Pig:
13.0
nTPM
Mouse:
4.3
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
48.5
Sample id
Sample description
nTPM
Hippocampus nTPM: 41.9
1911
Female, age 85, right hemisphere
46.2
2529
Female, age 94, left hemisphere
45.3
2148
Female, age 91, left hemisphere
41.7
1855
Male, age 76, left hemisphere
40.8
1898
Male, age 80, left hemisphere
39.3
2621
Female, age 67, right hemisphere
38.0
Dentate gyrus nTPM: 33.7
2246
Female, age 85, both hemispheres
46.2
2528
Male, age 74, both hemispheres
41.8
1927
Female, age 91, both hemispheres
35.6
2010
Male, age 76, both hemispheres
32.4
2530
Female, age 94, left hemisphere
25.8
2531
Female, age 67, both hemispheres
20.3
Entorhinal gyrus nTPM: 40.4
2686
Male, age 75, right hemisphere
51.6
2613
Female, age 94, left hemisphere
41.7
2117
Female, age 91, right hemisphere
40.3
2764
Male, age 80, right hemisphere
39.8
1938
Female, age 85, left hemisphere
39.7
1787
Male, age 76, left hemisphere
29.5
Hippocampus, CA1 nTPM: 48.5
2749
Male, age 80, left hemisphere
75.5
2527
Male, age 74, both hemispheres
55.2
1873
Female, age 85, both hemispheres
43.7
2082
Female, age 91, right hemisphere
42.8
2045
Male, age 76, both hemispheres
25.1
Hippocampus, CA2 nTPM: 38.9
1963
Male, age 80, left hemisphere
50.8
2249
Male, age 74, both hemispheres
44.0
2113
Female, age 91, right hemisphere
43.5
1892
Male, age 76, both hemispheres
32.4
2437
Female, age 94, left hemisphere
31.5
1940
Female, age 85, left hemisphere
31.1
Hippocampus, CA3 nTPM: 39.9
2137
Male, age 80, left hemisphere
66.7
2436
Male, age 74, both hemispheres
44.9
1891
Female, age 85, both hemispheres
37.6
1867
Male, age 76, both hemispheres
34.3
2861
Female, age 67, both hemispheres
32.0
2620
Female, age 94, left hemisphere
24.2
Parahippocampal cortex nTPM: 37.6
2692
Male, age 74, both hemispheres
54.0
2622
Female, age 67, right hemisphere
47.4
2250
Female, age 67, right hemisphere
41.3
1850
Male, age 80, both hemispheres
37.1
2245
Female, age 91, both hemispheres
35.8
2693
Female, age 94, both hemispheres
35.7
1933
Female, age 85, both hemispheres
35.1
2018
Male, age 76, both hemispheres
33.1
1990
Female, age 91, right hemisphere
33.0
2532
Female, age 72, left hemisphere
23.5
Perirhinal gyrus nTPM: 30.9
2521
Female, age 94, right hemisphere
35.3
1847
Female, age 91, left hemisphere
34.9
2848
Male, age 76, right hemisphere
32.2
1794
Male, age 80, right hemisphere
28.6
1841
Female, age 85, right hemisphere
23.7
Subiculum nTPM: 42.2
2619
Male, age 74, both hemispheres
48.2
2729
Female, age 91, right hemisphere
44.2
1901
Male, age 80, both hemispheres
43.5
2438
Female, age 94, left hemisphere
42.5
2042
Male, age 76, both hemispheres
38.5
1977
Female, age 85, both hemispheres
36.2
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
17.2
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 17.2
Samples: 197
Max nTPM: 41.5
Min nTPM: 2.5
Std nTPM: 8.3
Median nTPM: 15.4
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
41.5
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
39.5
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
37.6
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
37.3
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
36.4
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
34.6
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
33.7
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
32.2
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
32.2
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
32.0
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
31.1
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
31.1
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
30.5
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
29.7
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
29.4
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
29.3
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
29.2
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
29.2
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
29.1
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
29.0
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
28.8
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
28.8
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
28.8
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
28.5
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
28.3
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
28.3
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
27.9
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
27.6
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
27.5
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
27.0
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
26.7
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
26.5
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
26.4
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
26.0
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
25.9
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
25.7
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
25.5
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
25.4
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
25.3
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
25.1
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
25.0
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
25.0
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
24.9
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
24.9
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
24.5
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
24.4
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
24.3
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
24.3
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
24.0
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
23.2
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
23.2
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
22.7
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
22.6
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
22.2
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
22.1
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
22.1
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
22.1
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
22.1
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
22.1
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
21.9
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
21.6
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
21.6
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
21.6
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
21.1
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
21.1
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
21.0
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
20.5
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
20.4
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
20.3
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
20.0
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
19.6
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
19.3
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
19.2
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
18.9
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
18.8
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
18.7
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
18.7
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
18.7
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
18.5
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
18.1
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
18.0
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
17.8
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
17.7
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
17.5
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
17.2
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
17.0
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
16.9
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
16.7
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
16.6
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
16.5
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
16.5
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
16.4
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
16.3
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
16.3
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
15.7
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
15.5
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
15.4
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
15.4
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
15.4
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
15.3
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
15.3
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
15.2
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
15.2
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
15.1
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
15.0
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
15.0
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
15.0
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
14.9
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
14.9
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
14.8
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
14.8
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
14.8
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
14.4
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
14.2
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
14.2
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
14.1
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
14.0
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
14.0
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
13.8
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
13.7
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
13.7
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
13.3
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
13.3
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
13.2
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
12.9
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
12.8
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
12.8
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
12.7
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
12.6
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
12.5
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
12.5
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
12.5
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
12.5
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
12.4
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
12.2
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
11.9
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
11.9
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
11.8
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
11.7
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
11.7
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
11.7
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
11.6
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
11.3
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
11.2
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
11.0
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
10.9
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
10.6
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
10.6
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
10.5
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
10.4
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
10.4
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
10.3
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
10.3
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
10.3
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
10.2
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
10.2
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
10.1
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
9.9
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
9.9
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
9.9
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
9.8
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
9.5
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
9.4
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
9.3
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
9.0
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
8.9
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
8.8
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
8.7
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
8.3
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
8.2
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
8.0
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
8.0
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
7.9
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
7.9
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
7.7
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
7.7
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
7.6
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
7.5
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
7.3
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
6.6
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
6.4
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
6.2
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
6.2
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
6.2
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
5.8
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
5.6
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
5.5
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
5.1
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
4.7
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
4.0
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
3.9
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
3.9
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
3.3
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
3.1
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
3.0
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
2.6
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
2.5
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.