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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
24.4
nTPM
HPA Brain:
14.2
nTPM
GTEx:
24.4
nTPM
FANTOM5:
11.1
Scaled Tags Per Million
Pig:
2.5
nTPM
Mouse:
1.0
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
14.2
Sample id
Sample description
nTPM
Hippocampus nTPM: 11.3
1898
Male, age 80, left hemisphere
13.0
2148
Female, age 91, left hemisphere
11.7
2529
Female, age 94, left hemisphere
11.2
2621
Female, age 67, right hemisphere
11.1
1911
Female, age 85, right hemisphere
10.4
1855
Male, age 76, left hemisphere
10.2
Dentate gyrus nTPM: 12.3
2010
Male, age 76, both hemispheres
16.1
2528
Male, age 74, both hemispheres
13.5
1927
Female, age 91, both hemispheres
11.9
2530
Female, age 94, left hemisphere
11.8
2246
Female, age 85, both hemispheres
10.6
2531
Female, age 67, both hemispheres
10.2
Entorhinal gyrus nTPM: 13.4
2686
Male, age 75, right hemisphere
14.4
1787
Male, age 76, left hemisphere
14.0
2764
Male, age 80, right hemisphere
13.8
2613
Female, age 94, left hemisphere
13.4
2117
Female, age 91, right hemisphere
13.2
1938
Female, age 85, left hemisphere
11.7
Hippocampus, CA1 nTPM: 12.3
1873
Female, age 85, both hemispheres
12.5
2082
Female, age 91, right hemisphere
12.5
2749
Male, age 80, left hemisphere
12.5
2527
Male, age 74, both hemispheres
12.1
2045
Male, age 76, both hemispheres
12.0
Hippocampus, CA2 nTPM: 13.1
1892
Male, age 76, both hemispheres
16.4
2249
Male, age 74, both hemispheres
14.9
1940
Female, age 85, left hemisphere
13.7
2113
Female, age 91, right hemisphere
11.7
1963
Male, age 80, left hemisphere
11.1
2437
Female, age 94, left hemisphere
10.8
Hippocampus, CA3 nTPM: 14.2
2436
Male, age 74, both hemispheres
18.9
1867
Male, age 76, both hemispheres
15.1
1891
Female, age 85, both hemispheres
14.6
2620
Female, age 94, left hemisphere
13.9
2137
Male, age 80, left hemisphere
12.4
2861
Female, age 67, both hemispheres
10.5
Parahippocampal cortex nTPM: 14.0
1850
Male, age 80, both hemispheres
15.4
2532
Female, age 72, left hemisphere
15.4
2693
Female, age 94, both hemispheres
14.9
1990
Female, age 91, right hemisphere
14.3
2692
Male, age 74, both hemispheres
14.2
2622
Female, age 67, right hemisphere
14.1
2018
Male, age 76, both hemispheres
13.4
2250
Female, age 67, right hemisphere
13.4
1933
Female, age 85, both hemispheres
12.4
2245
Female, age 91, both hemispheres
12.3
Perirhinal gyrus nTPM: 13.5
2848
Male, age 76, right hemisphere
15.0
1841
Female, age 85, right hemisphere
14.8
1794
Male, age 80, right hemisphere
13.4
1847
Female, age 91, left hemisphere
12.7
2521
Female, age 94, right hemisphere
11.5
Subiculum nTPM: 13.9
2042
Male, age 76, both hemispheres
17.6
2619
Male, age 74, both hemispheres
14.9
1901
Male, age 80, both hemispheres
14.6
2729
Female, age 91, right hemisphere
13.0
1977
Female, age 85, both hemispheres
12.2
2438
Female, age 94, left hemisphere
11.3
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
24.4
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 24.4
Samples: 197
Max nTPM: 73.6
Min nTPM: 12.9
Std nTPM: 6.4
Median nTPM: 23.7
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
73.6
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
47.7
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
43.6
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
41.6
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
39.3
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
36.8
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
36.7
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
35.1
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
34.4
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
34.3
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
34.2
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
33.7
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
32.8
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
32.6
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
32.5
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
31.6
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
31.3
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
30.8
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
30.8
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
30.6
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
30.6
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
30.2
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
30.2
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
29.8
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
29.6
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
29.3
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
29.2
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
29.2
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
29.1
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
29.0
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
29.0
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
28.7
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
28.4
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
28.4
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
28.2
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
28.0
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
27.8
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
27.6
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
27.6
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
27.6
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
27.5
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
27.5
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
27.4
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
27.4
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
27.3
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
27.2
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
27.1
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
27.1
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
27.1
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
27.0
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
27.0
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
26.9
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
26.9
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
26.8
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
26.7
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
26.6
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
26.6
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
26.5
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
26.5
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
26.5
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
26.4
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
26.4
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
26.4
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
26.3
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
26.3
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
26.2
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
25.8
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
25.8
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
25.8
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
25.7
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
25.5
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
25.5
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
25.4
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
25.4
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
25.4
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
25.4
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
25.3
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
25.1
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
24.9
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
24.9
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
24.7
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
24.7
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
24.4
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
24.4
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
24.4
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
24.3
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
24.2
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
24.2
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
24.1
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
24.1
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
24.0
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
24.0
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
24.0
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
24.0
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
24.0
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
23.9
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
23.7
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
23.7
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
23.7
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
23.7
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
23.7
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
23.6
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
23.5
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
23.4
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
23.4
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
23.4
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
23.3
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
23.1
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
23.1
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
23.0
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
23.0
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
23.0
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
23.0
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
22.9
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
22.9
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
22.7
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
22.7
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
22.7
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
22.6
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
22.6
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
22.6
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
22.5
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
22.5
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
22.5
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
22.3
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
22.2
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
22.2
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
22.1
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
22.0
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
22.0
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
22.0
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
21.9
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
21.8
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
21.4
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
21.3
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
21.3
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
21.3
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
21.2
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
21.0
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
21.0
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
21.0
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
20.9
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
20.9
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
20.8
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
20.8
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
20.7
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
20.6
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
20.6
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
20.5
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
20.2
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
20.2
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
20.1
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
19.8
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
19.8
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
19.8
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
19.7
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
19.6
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
19.5
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
19.4
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
19.3
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
19.3
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
19.3
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
19.0
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
18.8
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
18.8
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
18.7
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
18.7
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
18.5
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
18.5
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
18.5
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
18.4
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
18.4
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
18.2
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
18.1
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
18.0
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
17.8
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
17.7
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
17.5
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
17.5
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
17.5
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
17.4
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
17.4
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
17.4
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
17.3
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
17.3
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
17.0
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
16.9
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
16.8
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
16.7
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
16.5
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
16.2
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
15.8
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
15.5
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
14.9
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
14.8
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
14.0
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
12.9
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.