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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
12.5
nTPM
HPA Brain:
20.2
nTPM
GTEx:
12.5
nTPM
FANTOM5:
13.2
Scaled Tags Per Million
Pig:
17.1
nTPM
Mouse:
31.8
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
20.2
Sample id
Sample description
nTPM
Hippocampus nTPM: 19.2
1898
Male, age 80, left hemisphere
22.2
1911
Female, age 85, right hemisphere
19.5
2621
Female, age 67, right hemisphere
19.4
1855
Male, age 76, left hemisphere
19.0
2148
Female, age 91, left hemisphere
18.1
2529
Female, age 94, left hemisphere
17.2
Dentate gyrus nTPM: 17.3
1927
Female, age 91, both hemispheres
20.8
2530
Female, age 94, left hemisphere
18.3
2010
Male, age 76, both hemispheres
17.9
2528
Male, age 74, both hemispheres
16.2
2246
Female, age 85, both hemispheres
16.0
2531
Female, age 67, both hemispheres
14.3
Entorhinal gyrus nTPM: 18.2
2613
Female, age 94, left hemisphere
22.4
2117
Female, age 91, right hemisphere
20.7
1938
Female, age 85, left hemisphere
19.6
2764
Male, age 80, right hemisphere
16.3
1787
Male, age 76, left hemisphere
15.9
2686
Male, age 75, right hemisphere
14.5
Hippocampus, CA1 nTPM: 20.2
2749
Male, age 80, left hemisphere
34.8
1873
Female, age 85, both hemispheres
18.8
2527
Male, age 74, both hemispheres
18.1
2082
Female, age 91, right hemisphere
14.6
2045
Male, age 76, both hemispheres
14.4
Hippocampus, CA2 nTPM: 18.2
1963
Male, age 80, left hemisphere
20.8
1940
Female, age 85, left hemisphere
20.2
2249
Male, age 74, both hemispheres
18.2
2113
Female, age 91, right hemisphere
16.8
2437
Female, age 94, left hemisphere
16.6
1892
Male, age 76, both hemispheres
16.4
Hippocampus, CA3 nTPM: 18.9
2137
Male, age 80, left hemisphere
31.0
2861
Female, age 67, both hemispheres
19.8
2436
Male, age 74, both hemispheres
17.3
1867
Male, age 76, both hemispheres
16.2
1891
Female, age 85, both hemispheres
15.3
2620
Female, age 94, left hemisphere
13.9
Parahippocampal cortex nTPM: 16.4
2245
Female, age 91, both hemispheres
20.6
2250
Female, age 67, right hemisphere
19.2
1933
Female, age 85, both hemispheres
18.1
2622
Female, age 67, right hemisphere
17.1
1990
Female, age 91, right hemisphere
16.6
2693
Female, age 94, both hemispheres
15.9
2018
Male, age 76, both hemispheres
15.6
1850
Male, age 80, both hemispheres
15.0
2532
Female, age 72, left hemisphere
13.6
2692
Male, age 74, both hemispheres
12.5
Perirhinal gyrus nTPM: 16.6
1841
Female, age 85, right hemisphere
19.1
1847
Female, age 91, left hemisphere
17.4
1794
Male, age 80, right hemisphere
17.1
2521
Female, age 94, right hemisphere
16.4
2848
Male, age 76, right hemisphere
13.0
Subiculum nTPM: 17.4
1901
Male, age 80, both hemispheres
22.3
2042
Male, age 76, both hemispheres
19.5
2729
Female, age 91, right hemisphere
18.5
1977
Female, age 85, both hemispheres
17.4
2619
Male, age 74, both hemispheres
13.7
2438
Female, age 94, left hemisphere
13.0
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
12.5
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 12.5
Samples: 197
Max nTPM: 36.6
Min nTPM: 5.8
Std nTPM: 3.9
Median nTPM: 11.7
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
36.6
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
21.0
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
20.8
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
20.6
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
20.6
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
20.6
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
20.6
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
20.5
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
20.3
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
20.0
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
19.5
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
19.3
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
19.1
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
18.8
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
18.7
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
18.6
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
18.3
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
17.9
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
17.7
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
17.6
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
17.6
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
17.4
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
17.3
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
17.2
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
17.2
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
17.2
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
17.1
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
17.1
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
17.0
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
16.7
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
16.7
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
16.7
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
16.6
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
16.5
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
16.4
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
16.4
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
16.3
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
16.1
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
15.9
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
15.7
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
15.6
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
15.5
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
15.5
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
15.3
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
15.1
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
14.9
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
14.9
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
14.8
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
14.8
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
14.7
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
14.5
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
13.9
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
13.7
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
13.7
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
13.6
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
13.6
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
13.6
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
13.5
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
13.5
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
13.5
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
13.5
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
13.5
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
13.5
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
13.3
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
13.2
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
13.2
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
13.2
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
13.2
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
13.0
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
13.0
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
13.0
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
13.0
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
13.0
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
12.9
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
12.9
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
12.9
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
12.8
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
12.8
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
12.8
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
12.8
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
12.8
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
12.7
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
12.6
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
12.5
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
12.3
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
12.3
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
12.2
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
12.2
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
12.1
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
12.1
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
12.1
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
12.1
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
12.1
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
12.0
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
12.0
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
11.9
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
11.9
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
11.8
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
11.7
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
11.6
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
11.6
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
11.6
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
11.6
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
11.6
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
11.6
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
11.5
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
11.5
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
11.5
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
11.4
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
11.4
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
11.4
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
11.4
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
11.3
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
11.3
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
11.2
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
11.2
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
11.0
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
11.0
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
11.0
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
11.0
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
10.9
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
10.9
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
10.9
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
10.9
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
10.8
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
10.8
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
10.8
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
10.7
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
10.7
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
10.7
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
10.7
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
10.6
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
10.5
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
10.4
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
10.4
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
10.4
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
10.4
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
10.3
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
10.3
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
10.3
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
10.2
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
10.0
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
10.0
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
10.0
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
10.0
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
9.9
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
9.8
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
9.8
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
9.7
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
9.5
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
9.4
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
9.4
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
9.4
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
9.4
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
9.3
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
9.3
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
9.2
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
9.1
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
9.1
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
9.1
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
9.1
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
9.0
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
9.0
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
9.0
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
9.0
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
9.0
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
9.0
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
8.8
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
8.8
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
8.6
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
8.6
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
8.5
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
8.4
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
8.4
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
8.3
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
8.2
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
8.0
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
7.9
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
7.9
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
7.9
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
7.9
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
7.9
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
7.9
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
7.7
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
7.7
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
7.7
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
7.5
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
7.1
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
7.1
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
7.1
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
7.0
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
6.8
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
6.7
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
6.4
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
5.8
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
5.8
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
5.8
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.