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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
18.2
nTPM
HPA Brain:
80.0
nTPM
GTEx:
18.2
nTPM
FANTOM5:
109.6
Scaled Tags Per Million
Pig:
103.6
nTPM
Mouse:
40.2
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
80.0
Sample id
Sample description
nTPM
Hippocampus nTPM: 72.2
2148
Female, age 91, left hemisphere
121.7
2621
Female, age 67, right hemisphere
96.9
1898
Male, age 80, left hemisphere
83.2
1855
Male, age 76, left hemisphere
66.1
1911
Female, age 85, right hemisphere
39.9
2529
Female, age 94, left hemisphere
25.6
Dentate gyrus nTPM: 72.4
1927
Female, age 91, both hemispheres
107.5
2530
Female, age 94, left hemisphere
78.9
2010
Male, age 76, both hemispheres
78.5
2531
Female, age 67, both hemispheres
75.3
2528
Male, age 74, both hemispheres
49.3
2246
Female, age 85, both hemispheres
44.8
Entorhinal gyrus nTPM: 46.4
2764
Male, age 80, right hemisphere
60.7
1938
Female, age 85, left hemisphere
48.5
2613
Female, age 94, left hemisphere
47.9
2686
Male, age 75, right hemisphere
45.6
1787
Male, age 76, left hemisphere
42.2
2117
Female, age 91, right hemisphere
33.8
Hippocampus, CA1 nTPM: 80.0
2749
Male, age 80, left hemisphere
185.8
1873
Female, age 85, both hemispheres
75.4
2527
Male, age 74, both hemispheres
57.5
2045
Male, age 76, both hemispheres
53.5
2082
Female, age 91, right hemisphere
27.9
Hippocampus, CA2 nTPM: 58.3
1963
Male, age 80, left hemisphere
128.7
1892
Male, age 76, both hemispheres
85.3
2249
Male, age 74, both hemispheres
40.3
2437
Female, age 94, left hemisphere
40.2
2113
Female, age 91, right hemisphere
32.3
1940
Female, age 85, left hemisphere
22.8
Hippocampus, CA3 nTPM: 65.2
2861
Female, age 67, both hemispheres
117.6
2137
Male, age 80, left hemisphere
106.4
2620
Female, age 94, left hemisphere
52.0
1867
Male, age 76, both hemispheres
46.5
2436
Male, age 74, both hemispheres
36.6
1891
Female, age 85, both hemispheres
31.8
Parahippocampal cortex nTPM: 44.7
2245
Female, age 91, both hemispheres
76.9
2532
Female, age 72, left hemisphere
53.5
2622
Female, age 67, right hemisphere
52.4
1850
Male, age 80, both hemispheres
46.9
1990
Female, age 91, right hemisphere
41.8
1933
Female, age 85, both hemispheres
39.9
2693
Female, age 94, both hemispheres
38.8
2250
Female, age 67, right hemisphere
38.2
2692
Male, age 74, both hemispheres
34.1
2018
Male, age 76, both hemispheres
24.3
Perirhinal gyrus nTPM: 44.9
1847
Female, age 91, left hemisphere
54.6
1794
Male, age 80, right hemisphere
51.9
2521
Female, age 94, right hemisphere
46.7
2848
Male, age 76, right hemisphere
37.0
1841
Female, age 85, right hemisphere
34.1
Subiculum nTPM: 40.5
1901
Male, age 80, both hemispheres
61.6
2042
Male, age 76, both hemispheres
44.9
2729
Female, age 91, right hemisphere
40.2
1977
Female, age 85, both hemispheres
35.4
2438
Female, age 94, left hemisphere
34.4
2619
Male, age 74, both hemispheres
26.5
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
18.2
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 18.2
Samples: 197
Max nTPM: 90.8
Min nTPM: 0.0
Std nTPM: 16.7
Median nTPM: 13.3
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
90.8
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
88.5
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
72.5
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
64.4
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
58.4
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
56.5
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
56.0
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
53.2
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
51.9
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
50.4
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
49.7
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
48.2
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
47.9
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
46.6
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
46.2
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
45.2
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
45.2
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
44.7
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
44.1
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
39.6
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
39.5
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
39.3
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
38.2
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
37.4
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
36.3
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
36.3
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
36.0
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
35.8
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
35.7
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
35.0
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
34.6
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
34.5
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
34.3
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
34.0
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
33.9
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
33.4
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
32.9
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
32.9
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
31.7
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
31.5
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
29.6
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
29.6
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
29.4
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
29.3
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
28.9
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
28.5
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
28.3
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
27.8
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
27.1
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
27.1
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
26.5
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
26.4
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
26.3
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
25.9
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
25.4
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
24.4
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
24.1
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
23.9
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
23.6
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
23.0
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
22.4
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
22.1
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
22.0
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
21.9
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
21.6
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
21.5
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
21.5
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
21.1
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
20.8
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
20.4
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
20.2
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
19.4
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
19.2
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
19.1
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
18.5
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
17.7
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
17.6
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
17.3
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
17.2
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
17.0
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
16.8
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
16.6
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
16.3
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
16.2
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
16.1
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
16.1
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
15.9
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
15.5
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
15.2
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
15.1
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
15.0
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
14.9
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
14.7
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
14.5
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
14.3
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
14.0
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
13.9
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
13.9
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
13.3
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
13.3
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
12.8
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
12.7
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
12.7
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
12.6
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
12.1
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
12.0
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
12.0
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
11.9
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
11.9
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
11.5
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
11.2
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
11.1
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
11.0
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
11.0
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
10.7
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
10.7
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
10.7
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
10.6
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
10.6
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
10.4
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
10.2
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
10.2
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
10.1
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
9.3
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
9.2
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
8.7
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
8.5
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
8.4
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
8.4
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
8.4
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
8.3
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
8.3
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
8.2
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
8.2
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
8.2
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
6.7
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
6.3
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
6.2
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
6.1
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
6.1
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
6.0
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
5.9
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
5.9
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
5.8
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
5.8
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
5.6
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
5.4
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
5.3
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
5.2
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
5.2
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
5.1
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
5.1
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
5.0
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
4.9
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
4.9
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
4.9
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
4.9
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
4.9
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
4.9
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
4.8
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
4.4
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
4.1
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
4.0
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
3.8
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
3.6
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
3.4
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
3.2
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
3.1
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
3.1
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
2.8
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
2.4
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
2.3
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
1.9
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
1.9
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
1.8
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
1.6
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
1.5
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
1.4
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
1.1
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
0.6
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
0.4
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
0.4
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
0.4
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
0.3
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
0.3
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
0.2
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
0.2
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
0.1
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
0.1
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
0.0
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
0.0
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
0.0
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
0.0
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
0.0
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
0.0
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
0.0
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
0.0
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.