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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
34.3
nTPM
HPA Brain:
82.6
nTPM
GTEx:
34.3
nTPM
FANTOM5:
71.7
Scaled Tags Per Million
Pig:
31.8
nTPM
Mouse:
12.9
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
82.6
Sample id
Sample description
nTPM
Hippocampus nTPM: 79.4
2529
Female, age 94, left hemisphere
98.6
2148
Female, age 91, left hemisphere
86.9
1898
Male, age 80, left hemisphere
79.8
1855
Male, age 76, left hemisphere
72.6
1911
Female, age 85, right hemisphere
70.5
2621
Female, age 67, right hemisphere
68.2
Dentate gyrus nTPM: 75.2
2528
Male, age 74, both hemispheres
93.6
2530
Female, age 94, left hemisphere
81.7
2531
Female, age 67, both hemispheres
74.7
2246
Female, age 85, both hemispheres
73.0
2010
Male, age 76, both hemispheres
67.3
1927
Female, age 91, both hemispheres
60.9
Entorhinal gyrus nTPM: 82.6
2117
Female, age 91, right hemisphere
123.4
2764
Male, age 80, right hemisphere
80.6
2686
Male, age 75, right hemisphere
80.2
1938
Female, age 85, left hemisphere
74.8
1787
Male, age 76, left hemisphere
71.9
2613
Female, age 94, left hemisphere
64.7
Hippocampus, CA1 nTPM: 72.0
2527
Male, age 74, both hemispheres
79.9
1873
Female, age 85, both hemispheres
75.6
2082
Female, age 91, right hemisphere
75.2
2749
Male, age 80, left hemisphere
68.3
2045
Male, age 76, both hemispheres
60.8
Hippocampus, CA2 nTPM: 74.4
2113
Female, age 91, right hemisphere
85.2
2437
Female, age 94, left hemisphere
80.5
1963
Male, age 80, left hemisphere
78.0
1892
Male, age 76, both hemispheres
70.8
1940
Female, age 85, left hemisphere
66.8
2249
Male, age 74, both hemispheres
65.2
Hippocampus, CA3 nTPM: 75.5
2861
Female, age 67, both hemispheres
110.4
1891
Female, age 85, both hemispheres
72.1
1867
Male, age 76, both hemispheres
71.5
2137
Male, age 80, left hemisphere
67.7
2620
Female, age 94, left hemisphere
67.1
2436
Male, age 74, both hemispheres
64.1
Parahippocampal cortex nTPM: 74.8
2692
Male, age 74, both hemispheres
95.8
2622
Female, age 67, right hemisphere
88.2
2245
Female, age 91, both hemispheres
81.4
1990
Female, age 91, right hemisphere
77.5
1933
Female, age 85, both hemispheres
75.0
2532
Female, age 72, left hemisphere
75.0
1850
Male, age 80, both hemispheres
67.3
2250
Female, age 67, right hemisphere
64.2
2018
Male, age 76, both hemispheres
63.3
2693
Female, age 94, both hemispheres
60.4
Perirhinal gyrus nTPM: 66.4
1847
Female, age 91, left hemisphere
77.3
1794
Male, age 80, right hemisphere
76.7
2521
Female, age 94, right hemisphere
61.2
1841
Female, age 85, right hemisphere
58.3
2848
Male, age 76, right hemisphere
58.2
Subiculum nTPM: 81.4
2729
Female, age 91, right hemisphere
100.9
2619
Male, age 74, both hemispheres
96.2
2438
Female, age 94, left hemisphere
79.3
1977
Female, age 85, both hemispheres
72.9
2042
Male, age 76, both hemispheres
70.5
1901
Male, age 80, both hemispheres
68.8
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
34.3
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 34.3
Samples: 197
Max nTPM: 76.9
Min nTPM: 15.0
Std nTPM: 11.3
Median nTPM: 31.7
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
76.9
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
71.8
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
64.5
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
64.1
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
63.1
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
62.5
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
58.0
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
57.6
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
57.1
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
56.3
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
54.9
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
54.7
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
54.6
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
54.5
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
54.2
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
54.1
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
52.9
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
52.6
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
51.8
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
50.5
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
50.2
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
49.3
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
47.7
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
47.2
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
47.2
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
46.7
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
46.6
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
46.5
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
46.3
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
46.1
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
46.1
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
45.7
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
45.6
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
44.9
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
44.9
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
44.6
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
44.2
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
43.9
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
43.6
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
43.6
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
42.5
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
42.3
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
42.3
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
42.2
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
41.9
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
41.5
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
41.3
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
40.9
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
40.6
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
39.7
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
39.7
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
39.6
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
39.2
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
39.2
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
39.1
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
39.0
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
38.5
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
38.5
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
38.4
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
38.3
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
38.2
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
38.0
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
37.7
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
37.6
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
37.5
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
37.1
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
37.1
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
37.0
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
36.9
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
36.3
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
36.0
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
35.7
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
35.6
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
35.6
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
35.3
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
35.3
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
34.8
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
34.8
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
34.6
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
34.4
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
34.3
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
34.3
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
34.1
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
34.0
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
33.8
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
33.4
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
33.3
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
33.2
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
32.6
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
32.6
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
32.5
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
32.3
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
32.2
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
31.9
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
31.9
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
31.7
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
31.7
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
31.7
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
31.7
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
31.7
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
31.7
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
31.5
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
31.4
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
31.4
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
31.3
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
31.3
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
31.2
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
31.0
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
31.0
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
30.9
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
30.9
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
30.8
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
30.8
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
30.8
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
30.8
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
30.5
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
30.5
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
30.1
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
30.0
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
29.9
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
29.7
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
29.7
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
29.7
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
29.6
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
29.5
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
29.4
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
29.1
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
28.8
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
28.4
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
28.4
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
28.2
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
27.9
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
27.9
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
27.9
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
27.8
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
27.8
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
27.6
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
27.4
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
27.4
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
27.3
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
27.1
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
27.0
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
26.9
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
26.9
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
26.8
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
26.8
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
26.8
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
26.7
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
26.5
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
26.4
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
26.4
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
26.3
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
25.9
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
25.9
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
25.9
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
25.7
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
25.7
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
25.6
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
25.4
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
25.0
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
24.4
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
24.2
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
24.1
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
24.0
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
23.7
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
23.6
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
23.3
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
23.2
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
23.1
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
23.0
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
23.0
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
23.0
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
23.0
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
23.0
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
22.8
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
22.7
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
22.2
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
22.0
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
22.0
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
21.5
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
21.4
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
21.2
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
20.9
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
20.8
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
19.6
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
19.5
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
19.5
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
19.3
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
18.8
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
18.4
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
17.2
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
16.4
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
15.9
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
15.6
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
15.4
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
15.0
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
15.0
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.