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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
11.1
nTPM
HPA Brain:
74.6
nTPM
GTEx:
11.1
nTPM
FANTOM5:
56.7
Scaled Tags Per Million
Pig:
36.4
nTPM
Mouse:
54.0
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
74.6
Sample id
Sample description
nTPM
Hippocampus nTPM: 49.5
2148
Female, age 91, left hemisphere
67.8
2621
Female, age 67, right hemisphere
57.5
1898
Male, age 80, left hemisphere
49.0
1855
Male, age 76, left hemisphere
47.6
1911
Female, age 85, right hemisphere
43.7
2529
Female, age 94, left hemisphere
31.1
Dentate gyrus nTPM: 53.9
2531
Female, age 67, both hemispheres
87.7
1927
Female, age 91, both hemispheres
77.7
2246
Female, age 85, both hemispheres
44.2
2010
Male, age 76, both hemispheres
43.1
2528
Male, age 74, both hemispheres
39.6
2530
Female, age 94, left hemisphere
31.3
Entorhinal gyrus nTPM: 42.7
2613
Female, age 94, left hemisphere
53.5
1938
Female, age 85, left hemisphere
47.4
2764
Male, age 80, right hemisphere
44.5
1787
Male, age 76, left hemisphere
38.6
2686
Male, age 75, right hemisphere
37.5
2117
Female, age 91, right hemisphere
34.5
Hippocampus, CA1 nTPM: 65.4
2749
Male, age 80, left hemisphere
130.1
1873
Female, age 85, both hemispheres
57.6
2045
Male, age 76, both hemispheres
56.7
2527
Male, age 74, both hemispheres
41.6
2082
Female, age 91, right hemisphere
41.2
Hippocampus, CA2 nTPM: 43.8
1963
Male, age 80, left hemisphere
68.0
1892
Male, age 76, both hemispheres
48.7
2113
Female, age 91, right hemisphere
39.5
2437
Female, age 94, left hemisphere
36.9
1940
Female, age 85, left hemisphere
35.5
2249
Male, age 74, both hemispheres
34.5
Hippocampus, CA3 nTPM: 74.6
2137
Male, age 80, left hemisphere
141.4
2620
Female, age 94, left hemisphere
90.7
2861
Female, age 67, both hemispheres
70.0
2436
Male, age 74, both hemispheres
54.8
1867
Male, age 76, both hemispheres
53.0
1891
Female, age 85, both hemispheres
37.5
Parahippocampal cortex nTPM: 45.3
2245
Female, age 91, both hemispheres
64.2
2693
Female, age 94, both hemispheres
59.3
1933
Female, age 85, both hemispheres
48.1
2250
Female, age 67, right hemisphere
47.6
1990
Female, age 91, right hemisphere
44.1
2018
Male, age 76, both hemispheres
42.6
2532
Female, age 72, left hemisphere
42.1
1850
Male, age 80, both hemispheres
41.8
2622
Female, age 67, right hemisphere
38.7
2692
Male, age 74, both hemispheres
25.1
Perirhinal gyrus nTPM: 54.1
2521
Female, age 94, right hemisphere
69.8
1847
Female, age 91, left hemisphere
66.9
1794
Male, age 80, right hemisphere
51.1
1841
Female, age 85, right hemisphere
45.4
2848
Male, age 76, right hemisphere
37.4
Subiculum nTPM: 39.9
1901
Male, age 80, both hemispheres
62.6
1977
Female, age 85, both hemispheres
46.5
2729
Female, age 91, right hemisphere
43.4
2619
Male, age 74, both hemispheres
34.0
2042
Male, age 76, both hemispheres
29.6
2438
Female, age 94, left hemisphere
23.0
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
11.1
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 11.1
Samples: 197
Max nTPM: 30.3
Min nTPM: 0.1
Std nTPM: 7.4
Median nTPM: 10.6
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
30.3
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
30.2
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
29.2
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
28.8
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
28.7
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
25.7
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
25.0
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
24.4
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
23.9
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
23.8
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
23.7
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
23.6
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
23.3
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
23.0
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
22.9
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
22.7
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
22.3
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
22.3
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
22.3
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
21.8
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
21.7
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
21.5
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
21.2
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
20.8
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
20.6
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
20.1
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
20.1
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
20.0
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
19.8
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
19.5
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
19.5
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
19.3
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
18.9
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
18.8
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
18.6
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
18.6
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
18.2
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
18.2
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
18.1
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
18.0
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
17.8
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
17.7
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
17.6
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
17.6
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
17.4
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
17.2
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
17.1
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
16.8
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
16.7
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
16.6
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
16.6
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
16.6
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
16.5
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
16.5
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
16.4
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
16.4
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
16.1
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
16.1
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
15.6
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
15.5
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
15.5
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
15.4
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
15.3
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
15.2
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
14.8
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
14.3
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
14.1
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
14.0
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
13.6
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
13.4
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
13.3
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
13.2
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
13.0
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
12.8
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
12.6
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
12.6
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
12.4
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
12.3
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
12.0
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
12.0
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
12.0
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
11.7
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
11.6
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
11.3
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
11.3
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
11.3
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
11.3
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
11.2
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
11.0
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
11.0
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
10.9
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
10.9
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
10.7
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
10.7
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
10.7
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
10.7
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
10.6
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
10.6
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
10.6
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
10.4
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
10.4
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
10.3
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
10.1
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
9.7
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
9.7
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
9.6
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
9.5
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
9.5
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
9.4
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
9.2
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
9.1
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
9.0
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
8.8
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
8.6
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
8.5
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
8.1
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
8.1
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
8.0
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
8.0
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
7.7
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
7.4
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
7.2
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
7.2
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
7.1
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
6.7
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
6.4
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
6.3
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
6.3
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
6.2
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
6.2
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
6.1
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
6.0
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
6.0
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
6.0
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
5.9
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
5.7
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
5.6
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
5.4
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
5.4
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
5.3
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
5.2
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
5.0
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
4.9
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
4.8
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
4.8
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
4.7
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
4.6
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
4.6
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
4.6
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
4.5
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
4.5
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
4.4
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
4.3
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
4.3
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
4.2
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
4.0
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
4.0
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
3.9
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
3.8
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
3.6
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
3.5
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
3.4
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
3.2
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
3.2
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
3.2
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
3.1
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
2.9
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
2.9
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
2.8
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
2.7
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
2.7
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
2.7
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
2.6
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
2.6
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
2.2
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
2.0
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
1.9
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
1.9
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
1.9
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
1.9
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
1.7
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
1.6
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
1.6
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
1.3
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
1.2
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
1.2
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
1.1
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
1.1
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
1.0
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
0.8
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
0.8
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
0.7
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
0.6
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
0.5
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
0.5
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
0.2
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
0.1
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.