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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
26.1
nTPM
HPA Brain:
62.4
nTPM
GTEx:
26.1
nTPM
FANTOM5:
59.1
Scaled Tags Per Million
Pig:
1361.3
nTPM
Mouse:
35.1
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
62.4
Sample id
Sample description
nTPM
Hippocampus nTPM: 37.5
1911
Female, age 85, right hemisphere
54.6
1898
Male, age 80, left hemisphere
40.9
2621
Female, age 67, right hemisphere
39.2
1855
Male, age 76, left hemisphere
38.9
2529
Female, age 94, left hemisphere
25.9
2148
Female, age 91, left hemisphere
25.6
Dentate gyrus nTPM: 24.7
2010
Male, age 76, both hemispheres
42.8
1927
Female, age 91, both hemispheres
40.7
2246
Female, age 85, both hemispheres
25.4
2530
Female, age 94, left hemisphere
14.8
2528
Male, age 74, both hemispheres
14.6
2531
Female, age 67, both hemispheres
10.1
Entorhinal gyrus nTPM: 12.2
2117
Female, age 91, right hemisphere
19.8
1938
Female, age 85, left hemisphere
14.5
2686
Male, age 75, right hemisphere
10.8
2613
Female, age 94, left hemisphere
9.7
1787
Male, age 76, left hemisphere
9.4
2764
Male, age 80, right hemisphere
9.1
Hippocampus, CA1 nTPM: 62.0
2749
Male, age 80, left hemisphere
196.9
2527
Male, age 74, both hemispheres
43.7
1873
Female, age 85, both hemispheres
31.0
2082
Female, age 91, right hemisphere
24.5
2045
Male, age 76, both hemispheres
13.9
Hippocampus, CA2 nTPM: 36.0
1963
Male, age 80, left hemisphere
63.2
1892
Male, age 76, both hemispheres
52.7
2249
Male, age 74, both hemispheres
32.8
2113
Female, age 91, right hemisphere
30.4
1940
Female, age 85, left hemisphere
18.6
2437
Female, age 94, left hemisphere
18.0
Hippocampus, CA3 nTPM: 62.4
2137
Male, age 80, left hemisphere
229.0
1867
Male, age 76, both hemispheres
55.8
1891
Female, age 85, both hemispheres
33.1
2436
Male, age 74, both hemispheres
31.2
2861
Female, age 67, both hemispheres
16.7
2620
Female, age 94, left hemisphere
8.9
Parahippocampal cortex nTPM: 18.8
2245
Female, age 91, both hemispheres
49.1
2018
Male, age 76, both hemispheres
26.2
2250
Female, age 67, right hemisphere
25.9
2692
Male, age 74, both hemispheres
17.4
1933
Female, age 85, both hemispheres
16.2
2622
Female, age 67, right hemisphere
12.8
1990
Female, age 91, right hemisphere
12.1
2693
Female, age 94, both hemispheres
10.5
1850
Male, age 80, both hemispheres
10.0
2532
Female, age 72, left hemisphere
8.5
Perirhinal gyrus nTPM: 10.9
1847
Female, age 91, left hemisphere
14.4
2521
Female, age 94, right hemisphere
11.9
1794
Male, age 80, right hemisphere
11.0
1841
Female, age 85, right hemisphere
8.8
2848
Male, age 76, right hemisphere
8.4
Subiculum nTPM: 43.1
1901
Male, age 80, both hemispheres
112.7
2042
Male, age 76, both hemispheres
59.7
2438
Female, age 94, left hemisphere
26.4
2619
Male, age 74, both hemispheres
23.1
2729
Female, age 91, right hemisphere
19.5
1977
Female, age 85, both hemispheres
17.5
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
26.1
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 26.1
Samples: 197
Max nTPM: 132.7
Min nTPM: 1.9
Std nTPM: 23.8
Median nTPM: 17.4
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
132.7
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
116.1
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
105.6
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
96.4
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
96.0
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
90.7
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
84.9
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
83.9
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
79.9
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
79.1
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
78.8
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
78.1
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
75.7
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
72.8
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
70.9
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
70.2
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
68.6
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
64.7
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
62.5
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
61.7
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
60.7
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
58.8
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
58.0
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
51.4
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
50.8
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
50.8
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
50.4
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
48.6
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
46.1
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
46.1
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
45.3
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
44.4
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
42.4
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
42.3
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
41.3
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
41.2
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
40.4
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
39.9
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
39.8
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
39.2
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
38.8
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
38.4
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
38.2
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
37.8
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
37.5
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
36.1
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
35.9
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
35.8
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
35.3
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
34.7
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
34.5
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
33.4
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
32.7
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
32.7
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
31.7
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
31.3
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
31.0
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
30.3
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
30.0
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
29.6
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
29.3
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
29.3
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
28.7
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
28.5
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
27.9
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
27.7
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
27.6
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
27.4
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
27.3
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
25.8
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
25.6
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
25.6
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
24.8
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
24.3
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
24.2
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
23.8
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
23.7
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
23.7
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
23.6
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
23.4
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
23.1
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
22.5
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
22.0
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
21.6
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
21.3
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
20.5
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
20.1
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
19.4
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
19.3
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
19.1
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
18.8
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
18.8
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
18.2
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
17.8
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
17.6
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
17.5
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
17.5
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
17.4
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
17.4
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
17.4
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
17.4
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
17.2
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
17.1
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
17.0
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
16.8
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
16.8
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
15.8
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
15.5
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
15.4
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
15.3
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
15.1
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
15.0
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
15.0
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
14.7
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
14.7
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
14.5
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
14.5
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
14.4
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
13.7
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
13.7
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
13.3
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
13.3
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
12.9
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
12.7
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
12.7
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
12.6
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
12.2
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
12.2
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
12.1
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
12.0
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
11.7
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
11.4
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
11.4
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
11.2
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
11.1
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
11.1
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
11.0
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
10.9
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
10.7
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
10.6
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
10.4
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
10.4
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
10.3
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
10.1
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
10.1
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
10.0
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
9.8
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
9.8
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
9.6
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
9.3
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
9.2
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
9.1
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
9.1
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
9.0
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
9.0
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
8.8
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
8.7
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
8.6
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
8.5
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
8.4
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
8.3
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
8.2
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
7.9
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
7.9
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
7.8
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
7.7
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
7.2
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
7.0
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
6.8
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
6.8
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
6.7
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
6.5
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
6.5
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
6.5
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
6.5
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
6.3
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
6.3
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
6.1
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
5.8
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
5.8
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
5.7
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
5.5
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
5.3
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
5.1
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
5.1
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
4.6
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
4.4
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
4.3
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
4.3
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
3.6
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
3.5
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
3.2
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
2.7
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
2.6
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
2.5
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
2.1
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
1.9
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.