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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
38.4
nTPM
HPA Brain:
46.5
nTPM
GTEx:
38.4
nTPM
FANTOM5:
7.4
Scaled Tags Per Million
Pig:
69.6
nTPM
Mouse:
44.8
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
46.5
Sample id
Sample description
nTPM
Hippocampus nTPM: 45.6
2621
Female, age 67, right hemisphere
61.1
2529
Female, age 94, left hemisphere
46.4
1855
Male, age 76, left hemisphere
45.3
2148
Female, age 91, left hemisphere
41.5
1898
Male, age 80, left hemisphere
41.0
1911
Female, age 85, right hemisphere
38.5
Dentate gyrus nTPM: 42.1
2531
Female, age 67, both hemispheres
66.2
1927
Female, age 91, both hemispheres
39.6
2528
Male, age 74, both hemispheres
38.1
2010
Male, age 76, both hemispheres
37.6
2246
Female, age 85, both hemispheres
36.3
2530
Female, age 94, left hemisphere
34.7
Entorhinal gyrus nTPM: 35.5
2686
Male, age 75, right hemisphere
42.8
2764
Male, age 80, right hemisphere
38.2
2613
Female, age 94, left hemisphere
37.6
2117
Female, age 91, right hemisphere
33.3
1787
Male, age 76, left hemisphere
31.9
1938
Female, age 85, left hemisphere
29.3
Hippocampus, CA1 nTPM: 46.5
2527
Male, age 74, both hemispheres
60.6
2749
Male, age 80, left hemisphere
55.5
2082
Female, age 91, right hemisphere
41.8
2045
Male, age 76, both hemispheres
38.4
1873
Female, age 85, both hemispheres
36.1
Hippocampus, CA2 nTPM: 38.7
1963
Male, age 80, left hemisphere
50.0
2113
Female, age 91, right hemisphere
45.0
2249
Male, age 74, both hemispheres
38.2
1892
Male, age 76, both hemispheres
37.2
2437
Female, age 94, left hemisphere
34.6
1940
Female, age 85, left hemisphere
27.4
Hippocampus, CA3 nTPM: 46.1
2861
Female, age 67, both hemispheres
73.0
2137
Male, age 80, left hemisphere
58.4
2620
Female, age 94, left hemisphere
51.6
2436
Male, age 74, both hemispheres
35.6
1867
Male, age 76, both hemispheres
33.0
1891
Female, age 85, both hemispheres
25.1
Parahippocampal cortex nTPM: 38.4
2245
Female, age 91, both hemispheres
63.1
2250
Female, age 67, right hemisphere
45.6
2622
Female, age 67, right hemisphere
43.0
2018
Male, age 76, both hemispheres
42.4
2693
Female, age 94, both hemispheres
39.5
2532
Female, age 72, left hemisphere
34.9
1933
Female, age 85, both hemispheres
30.9
1990
Female, age 91, right hemisphere
30.1
1850
Male, age 80, both hemispheres
27.1
2692
Male, age 74, both hemispheres
27.0
Perirhinal gyrus nTPM: 38.3
2521
Female, age 94, right hemisphere
45.1
1847
Female, age 91, left hemisphere
41.9
1794
Male, age 80, right hemisphere
40.7
1841
Female, age 85, right hemisphere
32.6
2848
Male, age 76, right hemisphere
31.3
Subiculum nTPM: 30.9
2729
Female, age 91, right hemisphere
35.7
2619
Male, age 74, both hemispheres
34.7
1977
Female, age 85, both hemispheres
32.8
1901
Male, age 80, both hemispheres
31.9
2042
Male, age 76, both hemispheres
26.3
2438
Female, age 94, left hemisphere
24.1
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
38.4
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 38.4
Samples: 197
Max nTPM: 161.0
Min nTPM: 4.2
Std nTPM: 21.2
Median nTPM: 36.9
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
161.0
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
93.0
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
91.0
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
85.3
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
85.2
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
84.4
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
84.2
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
82.8
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
82.0
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
78.5
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
77.2
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
75.4
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
71.7
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
70.8
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
70.5
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
70.5
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
70.4
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
69.4
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
68.4
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
68.4
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
65.6
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
64.7
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
64.4
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
62.6
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
60.1
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
58.4
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
57.4
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
56.9
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
56.1
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
54.9
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
54.7
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
54.4
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
54.1
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
53.9
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
53.8
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
52.3
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
51.8
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
51.3
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
50.9
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
50.7
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
50.4
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
50.3
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
49.5
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
49.4
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
49.3
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
49.3
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
49.2
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
48.8
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
48.6
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
48.5
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
48.1
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
47.7
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
47.2
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
47.2
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
46.8
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
46.6
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
46.6
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
46.3
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
46.1
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
45.9
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
45.9
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
45.8
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
45.7
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
45.7
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
45.7
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
45.7
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
45.4
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
45.3
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
44.4
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
44.3
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
44.2
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
43.6
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
43.5
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
42.9
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
42.8
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
42.8
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
42.4
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
42.2
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
41.9
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
41.5
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
41.4
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
41.2
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
41.1
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
40.8
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
40.7
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
40.3
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
40.2
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
39.6
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
39.5
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
39.4
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
38.4
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
38.2
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
37.3
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
37.2
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
37.2
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
37.2
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
37.1
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
37.0
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
36.9
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
36.7
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
36.6
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
36.5
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
36.3
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
36.3
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
36.2
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
35.9
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
35.1
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
34.8
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
34.6
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
34.4
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
34.3
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
34.1
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
34.0
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
33.9
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
33.3
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
32.9
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
32.2
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
31.5
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
31.4
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
31.4
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
31.3
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
31.3
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
30.6
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
30.4
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
30.2
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
29.5
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
29.4
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
29.2
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
28.9
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
28.8
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
28.5
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
28.4
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
28.1
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
27.6
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
27.5
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
27.3
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
26.8
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
26.6
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
26.3
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
25.8
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
25.8
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
25.7
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
25.6
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
25.6
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
25.5
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
25.4
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
24.6
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
24.5
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
24.2
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
24.0
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
22.4
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
22.1
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
22.1
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
22.0
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
21.9
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
21.7
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
21.6
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
20.7
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
20.0
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
19.4
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
18.6
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
18.0
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
17.7
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
17.5
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
17.0
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
16.9
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
16.5
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
15.7
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
15.2
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
15.2
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
15.1
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
14.7
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
14.6
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
14.5
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
14.5
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
14.0
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
13.9
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
13.6
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
13.0
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
12.9
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
11.3
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
11.0
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
10.6
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
10.2
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
10.1
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
9.5
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
8.8
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
8.7
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
8.1
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
7.5
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
7.5
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
7.2
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
7.0
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
6.3
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
5.8
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
4.4
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
4.2
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.