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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
4.2
nTPM
HPA Brain:
145.3
nTPM
GTEx:
4.2
nTPM
FANTOM5:
99.5
Scaled Tags Per Million
Pig:
68.7
nTPM
Mouse:
59.6
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
145.3
Sample id
Sample description
nTPM
Hippocampus nTPM: 91.8
2148
Female, age 91, left hemisphere
146.9
2621
Female, age 67, right hemisphere
96.3
1855
Male, age 76, left hemisphere
86.7
1898
Male, age 80, left hemisphere
83.8
2529
Female, age 94, left hemisphere
70.2
1911
Female, age 85, right hemisphere
67.2
Dentate gyrus nTPM: 120.6
2531
Female, age 67, both hemispheres
256.3
1927
Female, age 91, both hemispheres
137.4
2528
Male, age 74, both hemispheres
123.7
2246
Female, age 85, both hemispheres
89.5
2010
Male, age 76, both hemispheres
86.2
2530
Female, age 94, left hemisphere
30.4
Entorhinal gyrus nTPM: 43.5
2764
Male, age 80, right hemisphere
107.8
2117
Female, age 91, right hemisphere
44.7
2686
Male, age 75, right hemisphere
38.9
1787
Male, age 76, left hemisphere
26.3
1938
Female, age 85, left hemisphere
24.4
2613
Female, age 94, left hemisphere
18.8
Hippocampus, CA1 nTPM: 140.7
2045
Male, age 76, both hemispheres
244.0
2082
Female, age 91, right hemisphere
125.7
2749
Male, age 80, left hemisphere
125.1
2527
Male, age 74, both hemispheres
117.1
1873
Female, age 85, both hemispheres
91.4
Hippocampus, CA2 nTPM: 59.5
2113
Female, age 91, right hemisphere
88.8
2249
Male, age 74, both hemispheres
88.3
1892
Male, age 76, both hemispheres
80.6
1940
Female, age 85, left hemisphere
45.3
2437
Female, age 94, left hemisphere
33.4
1963
Male, age 80, left hemisphere
20.4
Hippocampus, CA3 nTPM: 145.3
2620
Female, age 94, left hemisphere
309.4
2861
Female, age 67, both hemispheres
201.6
1867
Male, age 76, both hemispheres
124.1
2137
Male, age 80, left hemisphere
80.4
1891
Female, age 85, both hemispheres
78.7
2436
Male, age 74, both hemispheres
77.7
Parahippocampal cortex nTPM: 72.0
2018
Male, age 76, both hemispheres
128.8
2250
Female, age 67, right hemisphere
100.6
2245
Female, age 91, both hemispheres
86.1
1850
Male, age 80, both hemispheres
81.3
2693
Female, age 94, both hemispheres
76.3
2532
Female, age 72, left hemisphere
76.2
1933
Female, age 85, both hemispheres
72.6
2622
Female, age 67, right hemisphere
65.4
1990
Female, age 91, right hemisphere
21.7
2692
Male, age 74, both hemispheres
10.7
Perirhinal gyrus nTPM: 93.5
2521
Female, age 94, right hemisphere
109.1
1794
Male, age 80, right hemisphere
101.6
2848
Male, age 76, right hemisphere
89.5
1847
Female, age 91, left hemisphere
86.3
1841
Female, age 85, right hemisphere
80.8
Subiculum nTPM: 62.6
2729
Female, age 91, right hemisphere
141.4
1977
Female, age 85, both hemispheres
59.3
1901
Male, age 80, both hemispheres
57.8
2042
Male, age 76, both hemispheres
47.6
2438
Female, age 94, left hemisphere
41.6
2619
Male, age 74, both hemispheres
27.7
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
4.2
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 4.2
Samples: 197
Max nTPM: 53.5
Min nTPM: 0.0
Std nTPM: 6.7
Median nTPM: 1.6
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
53.5
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
36.4
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
35.2
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
29.0
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
23.9
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
17.9
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
15.5
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
15.3
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
15.2
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
15.0
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
14.5
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
14.5
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
13.4
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
13.1
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
13.0
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
12.9
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
12.7
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
12.7
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
12.4
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
11.5
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
11.3
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
11.2
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
11.2
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
11.2
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
10.7
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
10.6
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
10.6
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
9.5
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
9.2
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
9.0
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
9.0
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
8.9
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
8.4
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
7.9
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
7.8
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
7.3
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
7.3
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
7.3
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
7.2
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
6.6
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
6.6
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
6.5
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
6.4
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
6.3
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
5.9
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
5.6
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
5.6
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
5.4
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
5.3
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
5.2
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
5.2
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
5.0
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
4.9
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
4.8
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
4.7
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
4.7
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
4.5
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
4.4
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
4.3
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
4.1
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
3.9
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
3.8
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
3.8
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
3.7
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
3.5
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
3.5
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
3.4
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
3.3
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
3.3
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
3.0
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
3.0
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
2.9
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
2.9
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
2.8
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
2.8
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
2.7
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
2.6
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
2.6
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
2.5
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
2.4
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
2.2
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
2.1
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
2.1
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
2.1
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
2.0
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
2.0
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
1.8
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
1.8
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
1.7
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
1.7
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
1.7
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
1.7
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
1.7
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
1.6
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
1.6
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
1.6
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
1.6
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
1.6
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
1.6
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
1.5
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
1.5
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
1.5
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
1.5
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
1.5
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
1.5
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
1.4
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
1.4
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
1.4
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
1.4
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
1.3
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
1.3
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
1.3
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
1.3
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
1.3
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
1.3
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
1.2
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
1.1
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
1.1
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
1.1
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
1.1
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
1.1
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
1.1
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
1.1
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
1.0
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
1.0
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
1.0
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
1.0
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
1.0
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
0.9
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
0.9
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
0.8
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
0.8
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
0.8
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
0.8
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
0.8
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
0.7
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
0.7
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
0.7
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
0.7
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
0.6
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
0.6
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
0.6
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
0.6
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
0.5
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
0.5
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
0.5
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
0.5
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
0.5
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
0.5
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
0.5
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
0.4
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
0.4
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
0.4
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
0.4
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
0.4
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
0.4
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
0.4
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
0.4
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
0.4
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
0.4
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
0.3
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
0.3
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
0.3
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
0.3
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
0.3
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
0.3
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
0.3
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
0.3
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
0.3
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
0.3
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
0.2
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
0.2
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
0.2
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
0.2
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
0.2
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
0.2
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
0.2
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
0.2
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
0.2
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
0.2
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
0.1
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
0.1
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
0.1
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
0.1
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
0.1
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
0.1
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
0.1
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
0.1
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
0.1
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
0.1
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
0.1
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
0.0
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
0.0
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
0.0
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
0.0
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
0.0
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
0.0
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.