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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
53.0
nTPM
HPA Brain:
53.4
nTPM
GTEx:
53.0
nTPM
FANTOM5:
25.5
Scaled Tags Per Million
Pig:
13.9
nTPM
Mouse:
0.9
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
53.4
Sample id
Sample description
nTPM
Hippocampus nTPM: 32.8
1911
Female, age 85, right hemisphere
43.5
2529
Female, age 94, left hemisphere
43.0
1898
Male, age 80, left hemisphere
32.2
2621
Female, age 67, right hemisphere
31.9
2148
Female, age 91, left hemisphere
28.1
1855
Male, age 76, left hemisphere
18.1
Dentate gyrus nTPM: 36.4
2530
Female, age 94, left hemisphere
72.2
2010
Male, age 76, both hemispheres
37.7
2528
Male, age 74, both hemispheres
33.0
2246
Female, age 85, both hemispheres
28.4
2531
Female, age 67, both hemispheres
28.0
1927
Female, age 91, both hemispheres
19.2
Entorhinal gyrus nTPM: 42.5
2117
Female, age 91, right hemisphere
82.8
2764
Male, age 80, right hemisphere
61.9
2686
Male, age 75, right hemisphere
30.0
1787
Male, age 76, left hemisphere
27.7
1938
Female, age 85, left hemisphere
26.9
2613
Female, age 94, left hemisphere
25.7
Hippocampus, CA1 nTPM: 21.8
2527
Male, age 74, both hemispheres
29.0
1873
Female, age 85, both hemispheres
26.1
2082
Female, age 91, right hemisphere
24.7
2749
Male, age 80, left hemisphere
22.7
2045
Male, age 76, both hemispheres
6.6
Hippocampus, CA2 nTPM: 35.5
1963
Male, age 80, left hemisphere
54.6
2249
Male, age 74, both hemispheres
40.2
2437
Female, age 94, left hemisphere
35.7
1940
Female, age 85, left hemisphere
34.7
2113
Female, age 91, right hemisphere
31.0
1892
Male, age 76, both hemispheres
16.7
Hippocampus, CA3 nTPM: 21.6
1891
Female, age 85, both hemispheres
48.9
1867
Male, age 76, both hemispheres
27.9
2436
Male, age 74, both hemispheres
17.4
2861
Female, age 67, both hemispheres
13.9
2137
Male, age 80, left hemisphere
12.1
2620
Female, age 94, left hemisphere
9.7
Parahippocampal cortex nTPM: 28.1
2532
Female, age 72, left hemisphere
49.5
1990
Female, age 91, right hemisphere
46.4
2692
Male, age 74, both hemispheres
39.8
2245
Female, age 91, both hemispheres
29.8
2018
Male, age 76, both hemispheres
28.2
1933
Female, age 85, both hemispheres
25.8
1850
Male, age 80, both hemispheres
24.9
2250
Female, age 67, right hemisphere
13.3
2693
Female, age 94, both hemispheres
12.0
2622
Female, age 67, right hemisphere
11.4
Perirhinal gyrus nTPM: 15.5
1794
Male, age 80, right hemisphere
18.5
2848
Male, age 76, right hemisphere
18.2
2521
Female, age 94, right hemisphere
16.1
1847
Female, age 91, left hemisphere
13.4
1841
Female, age 85, right hemisphere
11.1
Subiculum nTPM: 53.4
2042
Male, age 76, both hemispheres
79.7
1901
Male, age 80, both hemispheres
70.7
1977
Female, age 85, both hemispheres
54.9
2438
Female, age 94, left hemisphere
45.5
2619
Male, age 74, both hemispheres
35.6
2729
Female, age 91, right hemisphere
34.0
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
53.0
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 53.0
Samples: 197
Max nTPM: 210.4
Min nTPM: 3.9
Std nTPM: 38.2
Median nTPM: 43.2
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
210.4
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
188.9
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
188.8
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
180.7
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
180.2
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
175.6
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
136.3
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
129.2
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
127.9
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
126.9
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
117.9
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
116.3
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
114.9
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
113.5
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
112.6
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
110.0
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
109.2
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
108.2
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
107.6
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
107.4
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
107.4
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
105.2
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
104.8
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
98.9
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
95.5
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
93.3
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
92.8
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
92.5
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
92.4
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
92.3
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
91.8
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
90.3
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
90.0
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
88.3
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
87.0
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
86.7
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
82.8
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
80.7
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
78.9
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
78.8
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
78.4
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
77.3
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
77.2
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
75.5
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
74.9
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
74.9
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
69.7
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
67.9
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
66.6
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
65.9
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
64.6
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
64.2
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
63.9
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
63.5
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
63.5
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
63.4
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
63.0
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
62.5
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
61.9
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
61.9
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
61.8
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
60.3
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
58.6
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
58.1
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
57.6
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
56.9
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
56.9
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
56.0
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
55.9
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
55.3
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
54.5
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
54.0
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
53.4
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
52.7
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
52.0
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
51.8
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
51.7
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
51.3
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
50.5
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
50.2
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
49.9
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
49.8
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
49.4
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
49.4
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
49.4
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
49.2
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
47.4
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
47.3
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
47.3
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
47.3
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
46.6
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
45.9
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
45.9
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
43.7
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
43.6
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
43.6
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
43.5
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
43.2
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
43.2
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
42.0
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
41.6
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
41.6
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
41.3
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
40.7
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
40.6
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
40.6
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
40.5
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
39.8
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
39.3
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
38.4
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
38.2
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
38.0
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
37.7
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
37.6
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
37.0
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
35.3
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
35.3
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
34.8
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
34.7
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
34.5
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
34.3
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
34.0
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
33.9
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
33.2
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
32.3
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
32.2
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
31.6
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
31.0
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
30.8
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
30.8
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
29.7
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
29.7
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
29.6
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
29.5
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
29.5
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
29.1
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
29.0
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
28.9
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
28.4
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
28.3
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
28.2
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
27.8
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
27.7
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
27.5
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
27.4
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
27.2
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
27.1
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
27.1
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
27.1
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
26.6
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
26.2
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
26.0
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
25.8
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
25.3
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
24.5
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
24.2
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
23.8
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
22.6
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
22.5
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
21.8
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
21.7
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
21.5
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
21.1
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
20.6
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
20.3
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
19.7
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
19.7
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
19.4
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
19.3
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
18.5
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
18.2
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
18.2
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
18.1
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
18.0
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
17.7
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
17.5
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
17.2
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
16.3
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
15.7
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
15.2
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
14.9
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
14.7
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
13.9
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
13.7
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
13.5
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
13.4
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
11.6
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
11.2
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
10.9
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
10.9
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
9.2
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
9.1
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
8.9
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
7.4
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
6.7
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
4.9
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
3.9
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.