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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
42.3
nTPM
HPA Brain:
113.1
nTPM
GTEx:
42.3
nTPM
FANTOM5:
112.8
Scaled Tags Per Million
Pig:
59.4
nTPM
Mouse:
39.8
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
113.1
Sample id
Sample description
nTPM
Hippocampus nTPM: 86.3
2621
Female, age 67, right hemisphere
108.7
2148
Female, age 91, left hemisphere
98.3
1898
Male, age 80, left hemisphere
86.3
1855
Male, age 76, left hemisphere
81.4
1911
Female, age 85, right hemisphere
72.2
2529
Female, age 94, left hemisphere
70.8
Dentate gyrus nTPM: 85.6
2531
Female, age 67, both hemispheres
128.5
1927
Female, age 91, both hemispheres
98.7
2528
Male, age 74, both hemispheres
89.7
2246
Female, age 85, both hemispheres
70.1
2530
Female, age 94, left hemisphere
67.1
2010
Male, age 76, both hemispheres
59.3
Entorhinal gyrus nTPM: 110.0
2613
Female, age 94, left hemisphere
146.4
2686
Male, age 75, right hemisphere
144.7
2764
Male, age 80, right hemisphere
107.4
1938
Female, age 85, left hemisphere
93.4
1787
Male, age 76, left hemisphere
88.6
2117
Female, age 91, right hemisphere
79.3
Hippocampus, CA1 nTPM: 92.9
2527
Male, age 74, both hemispheres
115.8
2749
Male, age 80, left hemisphere
110.2
2082
Female, age 91, right hemisphere
94.2
1873
Female, age 85, both hemispheres
74.5
2045
Male, age 76, both hemispheres
69.9
Hippocampus, CA2 nTPM: 80.9
2437
Female, age 94, left hemisphere
95.0
1892
Male, age 76, both hemispheres
92.0
1963
Male, age 80, left hemisphere
84.0
2113
Female, age 91, right hemisphere
82.5
2249
Male, age 74, both hemispheres
79.9
1940
Female, age 85, left hemisphere
51.8
Hippocampus, CA3 nTPM: 98.3
2861
Female, age 67, both hemispheres
154.4
2620
Female, age 94, left hemisphere
116.9
2137
Male, age 80, left hemisphere
105.8
2436
Male, age 74, both hemispheres
84.1
1867
Male, age 76, both hemispheres
70.0
1891
Female, age 85, both hemispheres
58.8
Parahippocampal cortex nTPM: 105.9
2622
Female, age 67, right hemisphere
146.2
2693
Female, age 94, both hemispheres
137.1
2245
Female, age 91, both hemispheres
124.5
2250
Female, age 67, right hemisphere
109.6
2692
Male, age 74, both hemispheres
107.2
1933
Female, age 85, both hemispheres
94.5
1850
Male, age 80, both hemispheres
91.2
2532
Female, age 72, left hemisphere
86.8
1990
Female, age 91, right hemisphere
85.9
2018
Male, age 76, both hemispheres
75.7
Perirhinal gyrus nTPM: 113.1
1847
Female, age 91, left hemisphere
129.5
2521
Female, age 94, right hemisphere
118.4
2848
Male, age 76, right hemisphere
114.2
1794
Male, age 80, right hemisphere
105.3
1841
Female, age 85, right hemisphere
98.2
Subiculum nTPM: 75.3
2619
Male, age 74, both hemispheres
93.1
2438
Female, age 94, left hemisphere
83.0
2729
Female, age 91, right hemisphere
82.3
1977
Female, age 85, both hemispheres
72.3
1901
Male, age 80, both hemispheres
71.2
2042
Male, age 76, both hemispheres
50.3
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
42.3
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 42.3
Samples: 197
Max nTPM: 105.7
Min nTPM: 0.5
Std nTPM: 22.2
Median nTPM: 43.0
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
105.7
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
101.3
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
100.8
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
97.6
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
86.7
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
84.7
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
82.1
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
81.6
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
81.4
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
80.7
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
79.4
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
78.8
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
78.7
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
74.4
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
73.9
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
73.1
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
72.9
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
71.5
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
69.8
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
69.1
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
69.0
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
68.6
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
68.2
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
67.7
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
67.6
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
67.1
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
67.0
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
66.5
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
65.4
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
64.7
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
64.4
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
63.9
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
63.8
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
63.7
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
63.4
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
62.3
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
61.9
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
61.7
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
61.7
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
61.4
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
61.3
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
60.9
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
60.3
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
60.0
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
59.9
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
58.9
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
58.9
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
58.8
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
57.7
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
57.5
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
57.3
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
57.1
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
57.0
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
56.5
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
56.4
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
56.0
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
55.7
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
55.0
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
54.9
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
54.3
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
54.0
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
53.9
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
52.4
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
52.4
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
52.3
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
52.1
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
51.0
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
50.9
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
50.7
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
50.7
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
49.8
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
49.7
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
49.4
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
49.3
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
49.1
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
48.5
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
48.1
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
47.9
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
47.8
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
47.5
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
47.2
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
46.8
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
46.8
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
46.6
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
46.0
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
45.8
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
45.0
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
44.8
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
44.8
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
44.6
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
44.6
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
44.4
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
44.4
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
44.3
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
44.0
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
43.9
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
43.8
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
43.1
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
43.0
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
42.6
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
41.6
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
41.3
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
40.9
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
40.6
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
40.4
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
40.4
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
40.2
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
39.6
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
39.3
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
39.1
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
38.4
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
38.3
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
38.3
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
37.7
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
37.5
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
37.5
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
37.1
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
36.8
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
36.3
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
36.1
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
36.0
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
35.9
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
35.8
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
35.7
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
35.0
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
34.7
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
34.6
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
34.4
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
34.0
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
33.9
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
33.9
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
33.8
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
32.7
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
32.2
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
31.3
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
30.7
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
30.4
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
30.2
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
28.9
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
28.3
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
27.7
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
27.5
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
27.2
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
26.3
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
25.3
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
25.0
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
24.8
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
24.4
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
24.3
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
22.9
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
22.3
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
21.9
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
21.6
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
21.3
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
21.2
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
21.0
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
21.0
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
20.5
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
20.3
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
20.0
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
19.7
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
19.6
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
19.4
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
18.6
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
18.2
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
17.1
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
16.9
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
16.8
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
16.6
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
16.6
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
16.1
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
15.8
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
15.8
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
15.4
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
14.5
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
14.2
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
13.5
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
13.2
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
11.5
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
11.1
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
10.7
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
10.2
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
9.0
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
8.4
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
7.8
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
7.5
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
7.4
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
5.9
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
5.6
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
5.2
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
4.7
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
3.8
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
3.6
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
3.4
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
1.6
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
1.1
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
0.5
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.